0000001 - Document - Cover Page link:presentationLink link:calculationLink link:definitionLink 9952151 - Statement - Condensed Consolidated Balance Sheets link:presentationLink link:calculationLink link:definitionLink 9952152 - Statement - Condensed Consolidated Balance Sheets (Parenthetical) link:presentationLink link:calculationLink link:definitionLink 9952153 - Statement - Condensed Consolidated Statements of Loss and Comprehensive Loss link:presentationLink link:calculationLink link:definitionLink 9952154 - Statement - Condensed Consolidated Statements of Loss and Comprehensive Loss (Parenthetical) link:presentationLink link:calculationLink link:definitionLink 9952155 - Statement - Condensed Consolidated Statement of Changes in Stockholders' Equity link:presentationLink link:calculationLink link:definitionLink 9952156 - Statement - Condensed Consolidated Statements of Cash Flows link:presentationLink link:calculationLink link:definitionLink 9952157 - Disclosure - Nature of Operations link:presentationLink link:calculationLink link:definitionLink 9952158 - Disclosure - Summary of Significant Accounting Policies link:presentationLink link:calculationLink link:definitionLink 9952159 - Disclosure - Recent Accounting Pronouncements link:presentationLink link:calculationLink link:definitionLink 9952160 - Disclosure - Net Loss Per Share link:presentationLink link:calculationLink link:definitionLink 9952161 - Disclosure - Cash, Cash Equivalents and Marketable Securities link:presentationLink link:calculationLink link:definitionLink 9952162 - Disclosure - Goodwill link:presentationLink link:calculationLink link:definitionLink 9952163 - Disclosure - Royalty Revenue Monetization link:presentationLink link:calculationLink link:definitionLink 9952164 - Disclosure - Stockholders' Equity link:presentationLink link:calculationLink link:definitionLink 9952165 - Disclosure - Research, Collaboration and Licensing Agreements link:presentationLink link:calculationLink link:definitionLink 9952166 - Disclosure - Leases link:presentationLink link:calculationLink link:definitionLink 9952167 - Disclosure - Commitments and Contingencies link:presentationLink link:calculationLink link:definitionLink 9952168 - Disclosure - Business Segments link:presentationLink link:calculationLink link:definitionLink 9955511 - Disclosure - Summary of Significant Accounting Policies (Policies) link:presentationLink link:calculationLink link:definitionLink 9955512 - Disclosure - Net Loss Per Share (Tables) link:presentationLink link:calculationLink link:definitionLink 9955513 - Disclosure - Cash, Cash Equivalents and Marketable Securities (Tables) link:presentationLink link:calculationLink link:definitionLink 9955514 - Disclosure - Royalty Revenue Monetization (Tables) link:presentationLink link:calculationLink link:definitionLink 9955515 - Disclosure - Stockholders' Equity (Tables) link:presentationLink link:calculationLink link:definitionLink 9955516 - Disclosure - Research, Collaboration and Licensing Agreements (Tables) link:presentationLink link:calculationLink link:definitionLink 9955517 - Disclosure - Leases (Tables) link:presentationLink link:calculationLink link:definitionLink 9955518 - Disclosure - Business Segments (Tables) link:presentationLink link:calculationLink link:definitionLink 9955519 - Disclosure - Summary of Significant Accounting Policies (Details) link:presentationLink link:calculationLink link:definitionLink 9955520 - Disclosure - Net Loss Per Share (Details) link:presentationLink link:calculationLink link:definitionLink 9955521 - Disclosure - Cash, Cash Equivalents and Marketable Securities - Schedule of Marketable Securities (Details) link:presentationLink link:calculationLink link:definitionLink 9955522 - Disclosure - Cash, Cash Equivalents and Marketable Securities - Schedule of Cash, Cash Equivalents and Marketable Securities Measured at Fair Value on a Recurring Basis (Details) link:presentationLink link:calculationLink link:definitionLink 9955523 - Disclosure - Goodwill (Details) link:presentationLink link:calculationLink link:definitionLink 9955524 - Disclosure - Royalty Revenue Monetization - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 9955525 - Disclosure - Royalty Revenue Monetization - Schedule of Liability Related to Sale of Future Royalties (Details) link:presentationLink link:calculationLink link:definitionLink 9955526 - Disclosure - Stockholders' Equity - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 9955527 - Disclosure - Stockholders' Equity - Schedule of Repurchase Agreements (Details) link:presentationLink link:calculationLink link:definitionLink 9955528 - Disclosure - Stockholders' Equity - Schedule of RSU and PSU Activity (Details) link:presentationLink link:calculationLink link:definitionLink 9955529 - Disclosure - Stockholders' Equity - Schedule of Stock Options Granted (Details) link:presentationLink link:calculationLink link:definitionLink 9955530 - Disclosure - Stockholders' Equity - Schedule of Stock-Based Compensation Expense for Equity Classified Instruments (Details) link:presentationLink link:calculationLink link:definitionLink 9955531 - Disclosure - Stockholders' Equity - Schedule of Estimated Fair Value of Stock Options Granted and Weighted-Average Option Pricing Assumptions (Details) link:presentationLink link:calculationLink link:definitionLink 9955532 - Disclosure - Research, Collaboration and Licensing Agreements - Schedule of Revenue Recognized from Strategic Partnerships (Details) link:presentationLink link:calculationLink link:definitionLink 9955533 - Disclosure - Research, Collaboration and Licensing Agreements - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 9955534 - Disclosure - Leases - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 9955535 - Disclosure - Leases - Schedule of Balance Sheet Classification of Lease Liabilities (Details) link:presentationLink link:calculationLink link:definitionLink 9955536 - Disclosure - Leases - Schedule of Maturities of Operating Lease Liabilities (Details) link:presentationLink link:calculationLink link:definitionLink 9955536 - Disclosure - Leases - Schedule of Maturities of Operating Lease Liabilities (Details) link:presentationLink link:calculationLink link:definitionLink 9955537 - Disclosure - Leases - Schedule of Cost Components of Operating Leases (Details) link:presentationLink link:calculationLink link:definitionLink 9955538 - Disclosure - Business Segments - Narrative (Details) link:presentationLink link:calculationLink link:definitionLink 9955539 - Disclosure - Business Segments - Schedule of Segment Reporting Information, by Segment (Details) link:presentationLink link:calculationLink link:definitionLink